The valves of an engine are held closed by valve springs. When the cam turns, its lobe pushes the valve down against the spring and as the lobe passes the valve, the spring pushes the valve back up to close it. This works well until you reach very high redlines at which point something called 'hysteresis' happens to the valve spring. Basically the spring can't push the valve closed fast enough and the valve 'floats', unable to completely close before its opened by the cam again. This is bad.
So some engineers at Renault's F1 team came up with a solution: use pneumatics in place of valve springs. Pneumatics use air to move things and therefore do not suffer from hysteresis. Pressurized nitrogen is used in the F1 engines to push the valves back up. This allows for reliable valve operation well past 21k rpms. MotoGP motorcycles are the only other form of racing to use pneumatic valves.In F1, you must qualify and race the same engine in two consecutive races. This rule was designed to control costs so that the teams did not build 'grenade' engines that would only last 1 qualifying session or one race. In practice, teams are free to run of their engines and change at any time. If your designated race engine blows up in its first qualifying or first race, you must take a 10-spot grid penalty for the next race.
The engine is also a structural (or 'stressed') member of the car. This means that the engine literally holds the rear of the car to the monocoque! The front of the engine is bolted to the rear bulkhead of the monocoque, and the rear of the engine is bolted to the transmission. The rear suspension is mounted to the transmission (also a stressed member). Lotus was one of the first teams to embrace this concept, as you can see here on their 1967 car.
Using the engine and transmission as stressed members allows the monocoque to be shorter and eliminates extra weight. This is in stark contrast to a NASCAR car in which the engine and transmission are mounted within a tube frame, and the suspension members and bodywork are bolted to this frame.
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